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Gene Mapping is the graphic representation of the arrangement of a gene or a DNA sequence on a chromosome. Also known as linkage mapping, it is used to locate and identify the gene or group of genes that determines a particular inherited trait.

The [course_title] course teaches linkage disequilibrium mapping that helps you to analyse non-familial data. Basic concepts of genetic variations will also be discussed in the course.

Prior knowledge of statistical tests and estimation is required.


This course does not involve any written exams. Students need to answer 5 assignment questions to complete the course, the answers will be in the form of written work in pdf or word. Students can write the answers in their own time. Each answer needs to be 200 words (1 Page). Once the answers are submitted, the tutor will check and assess the work.


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Course Credit: Kyoto University

Course Curriculum

Module: 1
Contents of Four Sections 00:02:00
Structure of Section 1 00:04:00
Module: 2
Part 1: How to Use Linkage Disequilibrium for Gene Mapping 00:05:00
Part 2: How is LD Measured? 00:09:00
Part 3: Calculation of LD index 00:13:00
Part 4: r^2 and D’ (LD indices) 00:04:00
Part 5: Parameterization of Haplotype Frequency 00:10:00
Part 6: Genotype from Haplotype 00:06:00
Part 7: Genotype Calculation with R 00:03:00
Part 8: Short Summary of This Section 00:02:00
Module: 3
Part 1: 2×3 Table and Single SNP Association Tests 00:21:00
Part 2: 2×3 Data Simulation and SNP Tests with R 00:04:00
Module: 4
Part 1: GWAS and Uniform Distribution of p-values 00:12:00
Part 2: Minimum p-value in GWAS 00:14:00
Part 3: Bonferroni’s Correction 00:05:00
Part 4: Multiple-testing Correction for Mutually Dependent Tests 00:09:00
Module: 5
Part 1: Mutations and Genetic Drift 00:13:00
Part 2: Allele Frequency, Effect Size and Power 00:26:00
Message to Students Who Finish This Course 00:03:00
Submit Your Assignment 00:00:00
Certification 00:00:00

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